BTT 2.428: shift+wheel switches Spaces, no such gesture defined

I'm on old 2.428, 10.15.4.

For some reason, shift+scrollwheel switches Spaces, even when there's no such gesture set. I use and have defined cmd+ctrl+opt+arrows to do that in MacOS System Preferences and tilt left/right on the mouse to trigger that.

It's a BTT problem as when I close it, the issue stops and reappears after relaunching BTT. Any clue as to what this might be? Thanks!